The Sabertooths, Part 4: The First Sabertoothed Cats

The early history of the two major saber-cat tribes isn’t well understood yet. (Martin and others, 2011d; Werdelin and others)

Different paleontologists call the first sabertoothed cats by different names. (Antón; van den Hoek Ostende and others; Werdelin and others) The most controversial names are “Miomachairodus,” “Machairodus,” and “Amphimachairodus.”

“This discussion is far from settled, but at the very least shows that these forms grade into one another.”
— Werdelin and others

While professional fossil-cat herders continue to debate the details and slowly reach a consensus, I have chosen two specific sources to follow, even though they aren’t totally consistent:

  • Mauricio Antón calls those early sabertooths Macharoidus and describes Amphimachairodus as a separate group. (Antón)
  • Werdelin and others suggest that “Miomachairodus,” “Machairodus,” and “Amphimachairodus,” whatever their exact relationships to one another were, are the ancestors – somehow – of Homotherium (Werdelin and others), the scimitar-toothed sabercat.

Okay. Let’s move on to the Mediterranean region as it was in 12 Ma (million years ago), when a time traveler might have seen Machairodus there (at a safe distance) and mistaken it for one of today’s big cats.

Machairodus was generally built like a modern tiger, except for a very narrow face, a longer neck than any big cat has today, and a short tail. (Antón; Turner and Antón)

Machairodus also had saberteeth . . . and advanced ones at that. (Antón)

Experts have painstakingly pieced Machairodus together from fossil fragments found in a natural carnivore trap near Madrid.

Those sabers are impressive, but they look a little strange to paleontologists. Perhaps “out of place” is a better way to put it.

The saberteeth are so advanced, they might have belonged to Homotherium, but Machairodus doesn’t have the rest of a sabertoothed cat’s features, like massive cranial structures to support powerful jaw muscles, powerful forelimbs, and a short hindlegs. (Antón)

In fact, this very feline animal had long back legs that probably made it quite a good jumper. (Antón; Turner and Antón)

So our time traveler didn’t make it back to today because they saw a tiger-sized sabertoothed cat on the other side of a wide crevice and thought they were safe.

For paleontologists, the saberteeth show that the Homotheriini evolved saberteeth first and then worked out the rest of the cranioskeletal adaptations later. Scientists call this process mosaic evolution. LINK (Antón)

Even without other physical characteristics that would enhance the advantages of saberteeth, Machairodus thrived. (Antón)

Its earliest fossils come from Turkey and Africa. After that, these saber-cats spread out across both Eurasia and North America. (van den Hoek Ostende and others)

Machairodus was the first member of the cat family to reach the size of modern lions and tigers, and it didn’t stop there. One of its most successful species, M. aphanistus, was over 3 feet tall (100 cm) at the shoulder. (Antón)

Perhaps being big offset the lack of other sabertooth features.

Saberteeth allowed for a faster kill. (Antón; Turner and others) Machairodus might not have been able yet to tackle the plant-eating giants that thrived back then, its size meant that it could have taken and eaten prey as big as what today’s big cats can handle. (Deng and others)


This was quite an advantage in a highly competitive environment.

Over time, sabertooths became more efficient as they slowly developed all the other physical marks of an advanced sabertooth. (van den Hoek Ostende and others)

They were ready to go, once they had the whole “sabertooth complex.” (Antón)

This included, but wasn’t limited to, powerful forelegs and shoulders to overpower prey and a long, muscular neck to safely position the sabers for that killing bite. (Antón; Turner and others)

There were still a few limits, though.

Megaherbivores as huge as the junior mastodons that Homotherium would eventually tackle in the Pleistocene existed back in the Miocene (Antón; Agustí and Antón; Prothero, 2006), but they were probably safe from all of the early sabertooths. (Antón)

Machairodus followed smaller prey – zebra-sized migrating horses – out of central and western Asia into Europe, where it found lots of other suitable plant-eaters. (Agustí )

This sabertoothed cat coexisted with the barbourofelids (Agustí ; Antón) and held its own against Europe’s big amphicyonid beardogs and ursids (members of the bear family Ursidae). (Hunt, 1989)

Around 10 Ma, a more advanced sabertooth – Amphimachairodus – replaced Machairodus. (Antón)


That’s an Amphimachairodus skull (bottom) and drawings of the partial skull of Proailurus, the Dawn Cat (top) – the probable ancestor of Pseudaelurus and therefore of both modern cats and sabercats.

Some fifteen million years of evolution separate the two.

A-Mach was even bigger than Machairodus – almost 4 feet high (120 cm) at the shoulder – and its head was longer and narrower. (Turner and Antón)

This saber-cat also had a very muscular neck, as well as enormous dewclaws for holding onto prey animals like horses and antelope. (Antón)

However, A-Mach still overall looked like a modern cat. (Antón; Werdelin and others)

That is, it was still very primitive.

Today we tend to think the sabertooths were primitive because they went extinct. Actually, Homotherium and Smilodon were very advanced specialists. (Antón; Turner and Antón) It’s today’s cats that are the die-hard conservatives.

No cat owner will be surprised by this, of course, but it shows up not only in mood but also in the cat’s build.

Modern felines still have the general Pseudaelurus body plan. They have just changed the legs and feet a bit to make it easier to move on the ground. (Antón; Turner and Antón)

Sabercats were much more specialized, and they still managed to cover a lot of ground.

A “veritable freeway” for both predators and prey existed across the Bering land bridge between Eurasia and North America. (Prothero, 2006, including quote)

In addition, a tectonic collision between Africa and Eurasia had opened up routes between these two continents.

Primitive members of the elephant family, which would eventually evolve into Ice-Age mastodons and mammoths, ventured out of Africa into the wider world while Eurasian sabertooths followed prey into Africa’s mosaic landscapes of forest and wooded grasslands. (Agustí and Antón; Strömberg)

Fossils of these African sabertoothed cats from as far back as 10-11 Ma are still being excavated and studied. (Antón; Peigné and others)

Perhaps the most impressive to date is Machairodus kabir from Chad. This enormous saber-cat might have weighed almost half a metric ton! (Peigné and others)

That’s plenty big enough to take down plant-eaters weighing close to a ton. (Peigné and others)

Another sabertooth, with the jaw-breaking name of Lokotunjailurus, has been found in both Chad and Kenya. The size of a modern lioness, it goes back to about 7 Ma. (Antón; de Bonis and others)

At first, paleontologists thought this long-limbed, graceful sabertoothed cat was Machairodus. However, further study has shown that Lokotunjailurus was a separate species, with some similarities to Homotherium. (Antón; van den Hoek Ostende and others)

It’s too bad there are no common names for these beautiful animals. The scientific names are so long, especially in a family tree!

Some paleontologists (Werdelin and others) draw a very general family tree of the Homotheriini this way: Machairodus around 12 Ma, through Amphimachairodus at about 10 Ma, to Homotherium.

Perhaps Lokotunjailurus fits in there between Amphimachairodus and Homotherium. (van den Hoek Ostende and others)

Finally, according to phylogenetic computer analyses, Homotherium showed up in Europe around 6 Ma and then in North America two or three million years later. (Werdelin and others)

Its earliest fossils go back to 4 Ma – about when the oldest known “Felis” modern-cat ancestor lived in Greece. The sabercat remains were found in both Africa and the Ukraine, which means that Homotherium could have evolved in either Eurasia or Africa. (Antón and others; Werdelin and others)

The roots of Smilodon’s tribe

Many cat fossils are still buried by time, waiting to be discovered. (Johnson and others)

Smilodon’s ancestors are among the missing. (Antón; Werdelin and others) However, some possible early candidates have been recognized. There was Promegantereon, around 11 to 10 Ma. (Werdelin and others)

It was the size of a puma, with saberteeth that weren’t very long but were flat in cross-section, with a curve. (Salesa and others, 2010b)

Promegantereon was smaller than Machairodus, but it had some of the other sabertooth complex features, including powerful forelegs, a big dewclaw, and a narrow head. (Antón; Turner and Antón)

This saber-cat was still primitive, like Machairodus, but already it was built to wrestle down prey (Salesa and others, 2010a) – a trademark of the Smilodontini hunting style.

Even early on, strength mattered in this group.

Promegantereon hunted deer, antelope (Turner and Antón), and other prey in Eurasia and Africa from about 10 Ma to 8 Ma. Then leopard-sized Paramachaerodus showed up around 8 Ma and held the ambush-and-slash niche for another three million years. (Salesa and others, 2010b)

Paramachaerodus had moderately long, serrated sabers. Its other sabertooth features were a little more developed than Promegantereon’s. (Salesa and others, 2010b; Turner and others)

Smaller saber-cats like these probably coexisted with lion-sized Amphimachairodus by sharing resources in some way that’s not well understood yet. (Werdelin and others)

Perhaps they took refuge in trees, just as modern leopards do today in lion and tiger country. (Antón; Domingo and others)


Toward the end of the Miocene, Europe’s rich wildlife suffered a crisis that was probably related to global cooling. Barbourofelids and amphicyonid beardogs were among the casualties, but the Knife-Tooths came through it with just some reorganization. (Agustí ; Prothero, 2006; van den Hoek Ostende and others)

In the new Pliocene epoch, Paramachairodus was replaced by Megantereon – an undisputed member of the Smilodontini tribe. (Antón; Hunt, 2004; van den Hoek Ostende and others)


This jaguar-sized saber-cat got around – ranging across Eurasia, Africa, and North America – but apart from the spectacular skeleton displayed above, few of its fossils have been discovered yet. (Antón; Zhu and others)

As a result, no one is sure when or where Megantereon evolved. (Antón; Zhu and others)

This matters a lot to paleontologists, because the most iconic sabertooth of all – Smilodon – appeared after Megantereon’s arrival.

Everybody, laypeople as well as scientists, wants to know where Smilodon – the dirk-toothed sabercat from California’s La Brea asphalt seeps and the only fossil cat to have appeared in a hit movie (Ice Age) – came from.

Some experts think that Megantereon traveled into North America from Asia and evolved into Smilodon around 2.4 Ma. (Hunt, 2004)

There are other possibilities, for instance, that Smilodon and Megantereon were two groups, or that they and another saber-cat that its discoverers have named Rhizosmilodon shared a common ancestor. (Wallace and Hulbert)

It’s even possible that Megantereon was in Africa around the time of Lokotunjailurus, long before it appeared on other continents. (de Bonis and others)

Anyway, here we are at last, some two and a half million years before the present. The Miocene is far behind us, snow is beginning to fall, and vast areas of the continental polar regions are icing over. (Agustí and Antón)

The most advanced cats to ever walk this Earth – Homotherium, Megantereon, and Smilodon – are now well established and at their peak.

To be continued tomorrow


Featured image: Megantereon cultridens, Natural History Museum, Basel, Switzerland: Ghedoghedo. CC Att SA 4.0 International. Partial.

Machairodus aphanistus, Museo Arqueológico Regional de la Comunidad de Madrid (Alcalá de Heneres).

Combined image:

  • Top: Proailurus, Cope, E. D. 1880. On the extinct cats of America. American Naturalist. xiv (12):833-857, figure 1.
  • Bottom: Machairodus. Ghedoghedo. SEE PAGE CC att SA 3.0 unported, 2.5 Generic, 2.5 Generic, and 1.0 Generic.

Leopard in tree: Siddarth Maheshwari. CC BY-SA 3.0.

Megantereon, Natural History Museum, Basel, Switzerland: Ghedoghedo. CC Att SA 4.0 International.

Cited and uncited sources:

Agustí, J. and Antón, M. 2002. Mammoths, sabertooths, and hominids: 65 million years of mammalian evolution in Europe. New York and Chichester: Columbia University Press.

Akhmetiev, M. A., and Beniamovski, V. N. 2009. Paleogene floral assemblages around epicontinental seas and straits in Northern Central Eurasia: proxies for climatic and paleogeographic evolution. Geologica Acta. 7(12):297–309.

Andersson, K., and Werdelin, L. 2003. The evolution of cursorial carnivores in the Tertiary: implications of elbow joint morphology. Proceedings of the Royal Society London B (supplement). 270:S163-S165.

Antón, M. 2013. Sabertooth. Bloomington:Indiana University Press.

Antón, M.; Salesa, M. J.; Galobart, A.; and Tseng, Z. J. 2011. The Plio-Pleistocene scimitar-toothed felid genus Homotherium Fabrini, 1890 (Machairodontinae, Homotherini): diversity, paleogeography, and taxonomic implications. Quaternary Science Reviews. 96:259-268.

Averianov, A.; Obraztsova, E.; Danilov, I.; Skutschas, P.; and Jin, J. 2016. First nimravid skull from Asia. Nature, Scientific Reports. doi:10.1038/srep25812.

Barnett, R.; Barnes, I.; Phillips, M. J.; Martin, L. D.; Harington, C. R.; Leonard, J. A.; and Cooper, A. 2005. Evolution of the extinct sabretooths and the American cheetahlike cat. Current Biology, 15(15):R589-R590.

Barnosky, A. D. 2001. Distinguishing the effects of the Red Queen and Court Jester of Miocene mammal evolution in the northern Rocky Mountains. Journal of Vertebrate Paleontology. 21(1):172-185.

Benton, M. J.; Donoghue, P. C. J.; Asher, R. J.; Friedman, M.; Near, T. J.; and Vinther, J. 2015. Constraints on the timescale of animal evolutionary history. Palaeontologia Electronica, 18.1.1FC 1-106.

Benton, M. J. 2009. The Red Queen and the Court Jester: Species diversity and the role of biotic and abiotic factors through time. Science. 323(5915):728-732. Abstract.

Christiansen, P. (2013), Phylogeny of the sabertoothed felids (Carnivora: Felidae: Machairodontinae). Cladistics. 29: 543–559; abstract.
de Bonis, L.; Peigné, S.; Mackaye, H. T.; Likius, A.; Vignaud, P.; and Brunet, M. 2010. New sabre-toothed cats in the Late Miocene of Toros Menalla (Chad). Comptes Rendus Palevol. 9:221-227.

Bryant, H. N. 1991. Phylogenetic Relationships and Systematics of the Nimravidae (Carnivora). Journal of Mammalogy, 72(1):56-78.

Cain, M. L.; Bowman, W. D.; and Hacker, S. D. 2014. Ecology. Sunderland, Massachusetts: Sinauer Associates.

Cope, E. D. 1880. On the Extinct Cats of America. American Naturalist. xiv (12):833-857.

Deng, T.; Zhang, Y-X; Tseng, Z. J.; and Hou, S-K. 2016. A skull of Machairodus horribilis and new evidence for gigantism as a mode of mosaic evolution in machairodonts (Felidae, Carnivora). Vertebrata PalAsiatica. 54(4):302-318.

Domingo, M. S.; Domingo, L.; Badgley, C.; Sanisidro, O.; and Morales, J. 2013. Resource partitioning among top predators in a Miocene food web. Proceedings of the Royal Society B. 280:2012-2138.

Ezard, T. H. G.; Aze, T.; Pearson, P. N.; and Purvis, A. 2011. Interplay between changing climate and species’ ecology drives macroevolutionary dynamics. Science. 332(6027):349-351.

Flynn, J. J., and Galiano, H. 1982. Phylogeny of early Tertiary Carnivora with a description of a new species of Protictis from the middle Eocene of northwestern Wyoming. American Museum Novitates. 2725:1-64.

Gradstein, F. M.; Ogg, J. G.; and Hilgen, F. G. 2012. On the geologic time scale. Newsletters on Stratigraphy. 45(2):171-188.

Heske, E. J. Fall 2013 semester. Mammalogy 462, online class notes. Multiple lectures. . Last accessed December 11, 2015.

Holliday, J. A., and Steppan, S. J. 2004. Evolution of hypercarnivory: the effect of specialization on morphological and taxonomic diversity. Paleobiology. 30(1):108-128.

Hunt, Jr., R. M. 1989. Biogeography of the Order Carnivora, in Carnivore Behavior, Ecology, and Evolution, ed. J. L. Gittleman, J. L., 2:485–541 Ithaca, NY: Cornell University Press.

—. 2004. Global climate and the evolution of large mammalian carnivores during the later Cenozoic in North America. Bulletin of the American Museum of Natural History. 285:139-156.

Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; and Teeling, E. C. 2006. The Late Miocene Radiation of Modern Felidae: A Genetic Assessment. Science 311:73-77.

Kemp, T. S. 2006. The origin and early radiation of the therapsid mamma-like reptiles: a paleobiological hypothesis. 19:1231-1247. Journal compilation: European Society for Evolutionary Biology.

Kitchener, A. C., Van Valkenburgh, B., and Yamaguchi, N. 2010. Felid form and function, in Biology and Conservation of Wild Felids, ed. D. W. Macdonald and A. J. Loveridge, 83-106. Oxford: Oxford University Press.

Martin, L. D. 1980. Paper 287: Functional Morphology and the Evolution of Cats. Transactions of the Nebraska Academy of Sciences and Affiliated Societies. VIII:141-154.

Naples, V. L.; Martin, L. D.; and Babiarz, J. P. 2011 Introduction, The Other Saber-tooths: Scimitar-tooth cats of the Western Hemisphere, eds. Naples, V. L.; Martin, L. D.; and Babiarz, J. P. Baltimore: The Johns Hopkins University Press.

Martin, L. D.; Babiarz, J. P.; and Naples, V. L. 2011b. The osteology of a cookie-cutter cat, Xenosmilus hodsonae, in The Other Saber-tooths: Scimitar-tooth cats of the Western Hemisphere, eds. Naples, V. L.; Martin, L. D.; and Babiarz, J. P. 42-97. Baltimore: The Johns Hopkins University Press.

Martin, L. D.; Naples, V. L.; and Babiarz, J. P. 2011c. Revision of the New World Homotheriini, in The Other Saber-tooths: Scimitar-tooth cats of the Western Hemisphere, eds. Naples, V. L.; Martin, L. D.; and Babiarz, J. P. 185-199. Baltimore: The Johns Hopkins University Press.

Martin, L. D.; Babiarz, J. P.; and Naples, V. L. 2011d. A framework for the North American Homotheriini, in The Other Saber-tooths: Scimitar-tooth cats of the Western Hemisphere, eds. Naples, V. L.; Martin, L. D.; and Babiarz, J. P. 200-209. Baltimore: The Johns Hopkins University Press.

Morales, M. M., and Giannini, N. P. 2014. Pleistocene extinctions and the perceived morphofunctional structure of the neotropical felid ensemble. Journal of Mammal Evolution. 21:395-405.

O’Brien, S. J. and Johnson, W. E. 2007. The evolution of cats. Scientific American. 297 (1):68-75.

O’Brien, S. J.; Johnson, W.; Driscoll, C.; Pontius, J.; Pecon-Slattery, J.; and Menotti-Raymond, M. 2008. State of cat genomics. Trends in Genetics. 24(6):268-279.

Peigné, S.; de Bonis, L.; Likius, A.; Mackaye, H. T.; Vignaud, P.; and Brunet, M. 2005. A new machairodontine (Carnivora, Felidae) from the Late Miocene hominid locality of TM 266, Toros-Menalla, Chad. Comptes Rendus Palevol. 4:243-253.

Probiner, B. L. 2015. New actualistic data on the ecology and energetics of hominin scavenging opportunities. Journal of Human Evolution. 80:1-16.

Prothero, D. R. 2004. Did impacts, volcanic eruptions, or climate change affect mammalian evolution? Palaeogeography, Palaeoclimatology, Palaeoecology. 214:283-294.

—. 2006. After the Dinosaurs: The Age of Mammals. Bloomington and Indianapolis : Indiana University Press.

Prothero, D. R., and Heaton, T. H. 1996. Faunal stability during the Early Oligocene climatic crash. Palaeogeography, Palaeoclimatology, Palaeoecology. 127:257-283.

Radinsky, L. B. 1982. Evolution of the skull shape in carnivores. 3. The origin and early radiation of the modern carnivore families. Paleobiology. 8(3):177-195.

Ravelo, A. C.; Andreasen, D. H.; Lyle, M.; Olivarez Lyle, A.; and Wara, M. W. 2004. Regional climate shifts caused by the gradual global cooling in the Pliocene epoch. Nature. 429:263-267.

Rothschild, B. M., and Martin, L. D. 2011. Pathology in saber-tooth cats, in The Other Saber-tooths: Scimitar-tooth cats of the Western Hemisphere, eds. Naples, V. L.; Martin, L. D.; and Babiarz, J. P. 34-41. Baltimore: The Johns Hopkins University Press.

Rothwell, T. 2003. Phylogenetic Systematics of North American Pseudaelurus (Carnivora: Felidae). American Museum Novitates. 3403:1-64.

Salesa, M. J., Antón, M., Morales, J., and Peigné, S. 2011. Functional anatomy of the postcranial skeleton of Styriofelis lorteti (Carnivora, Felidae, Felinae) from the Middle Miocene (MN 6) locality of Sansan (Gers, France). Estudios Geológicos, 67(2):223-243.

Salesa, M. J.; Antón, M.; Turner, A.; and Morales, J. 2010a. Functional anatomy of the forelimb in Promegantereon ogygia (Felidae, Machairodontinae, Smilodontini) from the the late Miocene of Spain and the origins of the sabre-toothed felid model. Journal of Anatomy. 216:381-396.

Salesa, M. J.; Antón, M.; Turner, A.; Alcala, L.; Montoya, P.; and Morales, J. 2010b. Systematic revision of the late Miocene sabre-toothed felid Paramachaerodus in Spain. Palaeontology. 53(6):1369-1391.

Schellhorn, R. and M. Sammugaraja. 2015. Habitat adaptations in the felid forearm. Palaeontol Z confirm 89:261-269.

Smithsonian National Museum of Natural History. Geologic Time: The Story of a Changing Earth. Last accessed in summer of 2015.

Strömberg, C. A. E. 2011. Evolution of Grasses and Grassland Ecosystems. Annual Reviews of Earth and Planetary Science. 2011. 39:517-544.

Stuart, A. J. 2015. Late Quaternary megafaunal extinctions on the continents: a short review. Geological Journal. 50:338-363.

Sunquist, M. and Sunquist, F. 2002. Wild cats of the world. Chicago and London: University of Chicago Press.

Turner, A., and Antón, M. 1997. The Big Cats and Their Fossil Relatives: An Illustrated Guide to Their Evolution and Natural History. New York: Columbia University Press.

Turner, A., Antón, M., Salesa, M. J., and J. Morales, J. 2011. Changing ideas about the evolution and functional morphology of Machairodontine felids. Estudios Geológicos. 67(2): 255-276.

van den Hoek Ostende, L., Morlo, M., and Nagel, D. 2006. Fossils explained (52): Majestic killers: the sabretoothed cats. Geology Today. 22(4):150-157.

Van Valkenburgh, B. 1999. Major patterns in the history of carnivorous mammals. Annual Reviews of Earth and Planetary Science. 27:463-493.

—. 2007. Déjà vu: the evolution of feeding morphologies in the Carnivora. Integrative and Comparative Biology. 47 (1):147-163.

Van Valkenburgh, B., and Hertel, F. 1993. Tough times at La Brea: Tooth breakage in large carnivores of the late Pleistocene. Science, New Series. 261(5120):456-459.

Wallace, S. C., and Hulbert, Jr., R. C. 2013. A new machairodont from the Palmetto Fauna (Early Pliocene) of Florida, with comments on the origin of the Smilodontini (Mammalia, Carnivora, Felidae). PLoS ONE. 8(3): e56173.

Webb, S. D. 1987. Community patterns in extinct terrestrial vertebrates, in Organization of Communities Past and Present, ed. Gee, J. H. R. and Giller, P. S., 439-466. Oxford: Blackwell Scientific Publications.

Werdelin, L. 1989. Carnivoran Ecomorphology: A Phylogenetic Perspective, in Carnivore Behavior, Ecology, and Evolution, ed. Gittleman, J. L., 2:582-624. Ithaca, NY: Cornell University Press.

Werdelin, L.; Yamaguchi, N.; Johnson, W. E.; and O’Brien, S. J.. 2010. Phylogeny and evolution of cats (Felidae), in Biology and Conservation of Wild Felids, ed. D. W. Macdonald and A. J. Loveridge, 59-82. Oxford: Oxford University Press.

Wesley-Hunt, G. D. 2005. The Morphological Diversification of Carnivores in North America. Paleobiology, 31(1):35-55.

Wheeler, H. T. 2011. Experimental paleontology of the scimitar-tooth and dirk-tooth killing bites, in The Other Saber-tooths: Scimitar-tooth cats of the Western Hemisphere, eds. Naples, V. L.; Martin, L. D.; and Babiarz, J. P., 19-33. Baltimore: The Johns Hopkins University Press.

Yamaguchi, N., Driscoll, C. A., Kitchener, A. C., Ward, J. M., and Macdonald, D. W. 2004. Craniological differentiation between European wildcats (Felis silvestris silvestris), African wildcats (F. s. lybica) and Asian wildcats (F. s. ornata): implications for their evolution and conservation. Biological Journal of the Linnean Society. 83:47-63.

Zhu, M.; Schubert, B. W.; Liu, J.; and Wallace, J. C. 2014. A new record of the saber-toothed cat Megantereon (Felidae, Machairodontinae) from an Early Pleistocene Gigantopithecus fauna, Yanliang Cave, Fusui, Guangxi, South China. Quaternary International. 354:100-109.

This series on sabertooths was originally posted at my Robin Huntingdon blog about a year ago.

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